Sean and Whitey knew enough facts about the case to at least make a weak connection to Silent Ray and John O'Shea after listening to the tape.
- They assumed that Katie knew her killer because the old lady witness told them that she heard a "Hi" followed by the shot.
- They knew that the gun used to kill Katie was the gun (not just caliber or type, but THE gun) that Just Ray used in a liquor store robbery years ago based on the ballistic analysis.
- Katie was killed in the Harris' neighborhood, assumedly on her way to Brendan's house (they were leaving for Vegas the next day.)
- You can tell from the call that they were kids and they knew that the victim was a girl but were obviously trying to hide what they knew.
So all of this adds up to (in Sean and Whitey's mind): How can all the evidence point to Brendan and yet, based on the polygraph, he is not the killer? Answer: They were looking at the wrong brother. That at least warrants a conversation with the boys. The rest is explained by Qrious above.
As I stated in another post, from a writer's perspective, the call is simply the mechanism that allows the characters to break the case. Whether the call is a plausible case-breaker or whether the call itself is realistic seems to be the question. On the former, yes, on the latter, that remains open for debate.
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