doctor who - Why did a Dalek beg River Song for mercy?

The Dalek article on the Doctor Who wiki says:

The emotional capacity of Daleks was limited to largely negative
emotions (hate being chief among them), as "human" emotions such as
compassion and sentimentality were considered by them to be
weaknesses. However, they were capable of comprehending these
emotions, and often used them to exploit their enemies.

I'm not sure whether this idea was established in the classic series or if it's a new idea created by the revived post-2005 series. For example, in Genesis of the Daleks (1975), when the Dalek's creator Davros gave instructions to a scientist on the genetic modifications he wanted, the scientist objected "It'll mean creatures without conscience, no sense of right or wrong, no pity. They'll be without feeling or emotion", and Davros answered "Correct." So that seems to suggest that in this conception, they were entirely lacking in emotion. But in the 2005 episode Dalek, the Doctor said "The Dalek race was genetically engineered. Every single emotion was removed except hate." Similarly in the recent episode Into the Dalek (2014), the Doctor said "Dalek mutants are born hating. This is what stokes the fire, extinguishes even the tiniest glimmer of kindness or compassion."

I don't know of any dialogue that established they can feel fear or other "negative" emotions other than hate--the closest I could find was The Parting of the Ways (2005), where the Doctor said "Do you know what they call me in the ancient legends of the Dalek Homeworld? The Oncoming Storm. You might've removed all your emotions but I reckon right down deep in your DNA, there's one little spark left, and that's fear." That seems more like a speculation that something he knew for sure though. And as the wiki quote above says, they can at least understand human emotions well enough to try to manipulate us with them, and it's possible that's all the Dalek was doing when it yelled "mercy!"